1 Department of Genetics, Stanford University Medical School, Stanford, CA
2 Department of Molecular Genetics and Microbiology, University of Florida College of Medicine, Gainesville, FL
3 Departments of Molecular, Cellular, and Depelopmental Biology, and Ecology and Evolutionary Biology, University of Michigan, Ann Arbor, MI
4 Howard Hughes Medical Institute and Department of Biochemistry, Stanford University Medical School, Stanford, CA
5 Division of Biological Sciences, University of Montana, Missoula, Montana
+To whom correspondence should be addressed
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